B.A. in Biology, San Diego State University, San Diego, California, 1975
Ph.D. in Biology, University of California, San Diego, 1985
Thesis: Biochemical genetics of the circadian rhythm in Neurospora crassa: Studies on the cel strain.
Supervisor: Dr. Stuart Brody
Damon Runyon-Walter Winchell Cancer Fund Post-Doctoral Fellowship, Dept. of Biochemistry, University of Cambridge, 1985–1988
Wellcome Trust Post-Doctoral Research Fellowship, Dept. of Plant Sciences, University of Cambridge, 1990–1993 and 1993–1995
Temporary University Lecturer, Dept. of Zoology, University of Cambridge, 1995–1998
Visiting Scientist, Dept. of Plant Sciences, University of Cambridge, 1998–2002
Associate Professor, Dept. of Biology, York University, 2002–2019
Professor, Dept. of Biology, York University, 2019–present